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Transcription
Argopsis 2
while his figure of the crossection of a cephalodium suggests the
cerebriform type.
Zhalbruckner tried to reconcile the literature without a critical
study of material of either species, resulting in misstatements such
as citing A. megalospora from Kerguelen rather than from Campbell Island
and stating that the eight small spores coalesce to form a degenerate
brown mass.
Fortunately the type of Stereocaulon Argus Tav.l. and a duplicate
of the type collection of S. cymosum Crombie are present and fruiting
in the Taylor Herbarium at the Boston Society of Natural History. The
former specimen agrees with Th. Fries' description of Argopsis, the
second with the description of A. Friesiana Müll. Arg. Both are similar
in podetial characters and cephalodia to Stereocaulon ramulosum.
S.Argus has a hyaline paratheciun and hypothecium, asci apparently not
more than 2-spored, of which one early degenerates, leaving a single
large muriform spore very similar to those of Lopadium, remaining
hyaline until late, then becoming slightly yellowish brown, not the
deep brown muriform spore of Rhizocarpon. S. cymosum has a brownish
paratheciun (not carbonaceous in the apothecium sectioned), asci 4-8-
spored; ascospores with three transverse septa and an occasional longi-
tudinal or oblique septum, producing a few-celled muriform spore simi-
lar to those in Collema sect. Blennthallia. Were it not for the ex-
treme form of S.Argus, there is little to distinguish S.cymosum from
from Stereocaulon where the sterile material was referred. Until
more information is available, it is largely a matter of individual
opinion whether the generic limits should be circumscribed to include
only S.Argus or enlarged to include S. cymosum. I have adopted the
latter course. Both species seem to be endemic to small areas, but it
is possible that they have been referred to some variety of S. ramulosum