Intragroup Relations Dominance? Feeding of young? Acorn storage Defense of territory { cooperative aspects were routed intruders of each sex Ligons hyp. that nestlings are resources, in AW, if true, should help nestlings, juv, 1st year birds more because they might eventually move with them. Dom shouldn't show this bias. Kin sel, helpers should help both sexes equally. Alt. hypothesis that intermale competition balances out Scrub Jays, you might expect helpers to preferentially help younger , since they will stay longer, enlarge territory, etc. New groups should want more (both AW + SJ) as offspring perhaps?